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Tribbles are a Star Trek: TOS canon invention appearing in the fan-favorite episode The Trouble with Tribbles written by David Gerrold.
Gerrold, utilizing either vague Paramount ownership of the term, or by official permission, was an early marketer of toy tribbles at conventions and through mail order.
Fans also featured tribbles in their publications.
Gerrold and Tribbles
Gerrold himself had a complicated relationship with tribbles.
One of the best ST episodes is "The Trouble With Tribbles." David Gerrold tells an interesting story about it in his book of the same name (Ballantine publishers). He discovered the tribbles were so similar to Martian flat cats in a story by Robert Heinlein that he was afraid he would be accused of plagiarism. He wrote to Heinlein who replied that the basic plot had been used before his book too.
In 1969, Gerrold wrote: "... there are too many fans who have not realized that Star Trek is a broken promise, a vision of things that will never be. These are the rabid little trekkees who get excited at seeing a real tribble (damn it, it's only a powder puff!)." 
In either a crafty marketing move with Paramount, or a reflection of Paramount not covering all its bases (possibly both), David Gerrold temporarily got over this disdain and personally sold a lot of tribbles. From a 1972 ad in Star-Borne:
Tribbles are available in the following colors: soft russet, shaggy gray, fluffy white, velvet black, outrageous calico, shocking zebra, and miscellaneous. Pleas specify second and third choices when ordering. Sometimes we run out.Tribbles are the exclusive creation of David Gerrold, and are copyright in his name. They were created for STAR Trek, and have appeared in "THE TROUBLE WITH TRIBBLES" (live action) and the forthcoming "MORE TRIBBLES, MORE TROUBLES." (animated) .
In 1973, Gerrold combined his tribble sales with yet another fan campaign, this one for Star Trek: The Animated Series, and by proxy, the possibility of a Star Trek movie. From an extensive flyer by Gerrold:
We're trying to generate the biggest mail response in TV history [to encourage viewers for Star Trek: The Animated Series]. We need to reach 100,000 people or more... It's going to cost a lot of money, though, not to mention printing a paper costs. But we're not asking for contributions. We don't want to take your money without giving you fair value in return. That's why this letter is going out with an advertising packet. When you buy a tribble or a book, you're helping us cover the mailing cost to you, but to two other people as well.
In 1974, a fan, Cara Sherman wrote:
DAVID GERROLD ALL OVER THE PLACE, or The Man Who Solded Himself  ***David Gerrold is DAGE CO. (Obviously an acronym of DAvid GErrold), and is vending all kinds of books and other things. [snipped: list of books for sale] He is also still selling those... tribbles. Short haired, small (about the size of a hairy ping pong ball), large (about the size of a hairy tennis ball) and immense (about the size of a hairy soccer ball), 2, 4 and 8 dollars a piece; long-haired (same sizes but shaggier), 3 dollars each, 2 for 5, large 5 dollars each, 2 for 9, and immense 10 dollars each or 2 for 16.I'd tell you what I think, but greed speaks for itself.
In 2016, Gerrold explained that tribbles had their part of making him decide to stop going to cons:
I knew it was time for me to stop going to conventions when I showed up at one and there were thirty people selling Tribbles. You say to them, "You don't have the right," and they say, "Fuck you, you made enough money off Star Trek. Now it's my turn." This was the shift. In '72 or '73, you'd meet the fans, and they were grateful for the opportunity to meet people who worked on Star Trek. By '75 or '76, the attitude was "We own Star Trek now. The studio doesn't care. We do." 
Art and Ads Starring Tribbles!
from the third 1973 Equicon progress report, another ad for tribbles
some 1973 fan art portraying Gerrold and a little friend, artist is George Barr, illo is in the Equicon program book
a bit of David Gerrold from Satire Trek Comicordance, parody of the George Barr illo
a 1973 ad for Gerrold's wares, printed in the zine Monolith #3
David Gerrold, Do Tribbles Have Bones?
Not bones, but apparently a boner:
From CompuServe's Science Fiction Forum: #: 57397 S2/Star Trek / 06-Feb-86 17:08:07
Sb: #Tribble Bones
Fm: Diane Duane 70137,2260
To: [FB] All:The results of a phone call to David Gerrold: "Tribbles do not have bones. They have cartilage...Or rather," he said, "they have *one* bone." The requirements of a G-rated Sig make it impossible for me to tell you where that bone *is*. I'm allowed to say that the bone is located somewhere in the area of "the boy part of the tribble." David declined to tell how you tell "the boy part" of a tribble from "the girl part". He does, however, say that filleting a tribble is very easy. He also implied that it was fun for the tribble. I hope he didn't say it just for the shock value. best, DED.
Fan Publications with a Tribble Emphasis
- Tribbles are People Too!
- Tribble Dribbles
- The Tribble Owner's Guide and Manual
- The Tribble Teletype
- Tribble Trek
- Tribble Trouble
- ^ from a fan in Interstat #125 (March 1988)
- ^ from The Awful Offal
- ^ "If the animated Star Trek does well enough, perhaps Paramount can be convinced to bring back the live series -- or even a movie!
- ^ "The Man Who Solded Himself" is a pun on Gerrold's novel, "The Man Who Folded Himself."
- ^ from Romulan Wine #4, see that zine for D.C. Fontana's comments on bootleg material, fan purchases, legalities, and ethics
- ^ from "The Fifty Year Mission: The Complete, Uncensored, Unauthorized Oral History" by Edward Gross and Mark A. Altman, published in 2016 by St. Martin's Press